,this e,book was converted to ,,html1 with additional editing1 by ,jose ,menendez from the text edition produced by ,amy ,e4 ,zelmer4 ,the ,darwinian ,hypothesis99 by ,thomas ,h4 ,huxley ,,darwin ,,on ,,the ,,origin ,,of ,,species4 ,,there is a growing immensity in the speculations of science to which no human thing or thought at this day is comparable4 ,apart from the results which science brings us home and securely harvests1 there is an expansive force and latitude in its tentative efforts1 which lifts us out of ourselves and transfigures our mortality4 ,we may have a preference for moral themes1 like the ,homeric sage1 who had seen and known much3-- 8,cities of men ,and manners1 climates1 councils1 governments02 yet we must end by confession that 8,the windy ways of men ,are but dust which rises up ,and is lightly laid again10 in comparison with the work of nature1 to which science testifies1 but which has no boundaries in time or space to which science can approximate4 ,there is something altogether out of the reach of science1 and yet the compass of science is practically illimitable4 ,hence it is that from time to time we are startled and perplexed by theories which have no parallel in the contracted moral world2 for the generalizations of science sweep on in ever-widening circles1 and more aspiring flights1 through a limitless creation4 ,while astronomy1 with its telescope1 ranges beyond the known stars1 and physiology1 with its microscope1 is subdividing infinite minutiae1 we may expect that our historic centuries may be treated as inadequate counters in the history of the planet on which we are placed4 ,we must expect new conceptions of the nature and relations of its denizens1 as science acquires the materials for fresh generalizations2 nor have we occasion for alarms if a highly advanced knowledge1 like that of the eminent ,naturalist before us1 confronts us with an hypothesis as vast as it is novel4 ,this hypothesis may or may not be sustainable hereafter2 it may give way to something else1 and higher science may reverse what science has here built up with so much skill and patience1 but its sufficiency must be tried by the tests of science alone1 if we are to maintain our position as the heirs of ,bacon and the acquitters of ,galileo4 ,we must weigh this hypothesis strictly in the controversy which is coming1 by the only tests which are appropriate1 and by no others whatsoever4 ,the hypothesis to which we point1 and of which the present work of ,mr4 ,darwin is but the preliminary outline1 may be stated in his own language as follows3--8,species originated by means of natural selection1 or through the preservation of the favoured races in the struggle for life40 ,to render this thesis intelligible1 it is necessary to interpret its terms4 ,in the first place1 what is a species8 ,the question is a simple one1 but the right answer to it is hard to find1 even if we appeal to those who should know most about it4 ,it is all those animals or plants which have descended from a single pair of parents2 it is the smallest distinctly definable group of living organisms2 it is an eternal and immutable entity2 it is a mere abstraction of the human intellect having no existence in nature4 ,such are a few of the significations attached to this simple word which may be culled from authoritative sources2 and if1 leaving terms and theoretical subtleties aside1 we turn to facts and endeavour to gather a meaning for ourselves1 by studying the things to which1 in practice1 the name of species is applied1 it profits us little4 ,for practice varies as much as theory4 ,let the botanist or the zoologist examine and describe the productions of a country1 and one will pretty certainly disagree with the other as to the number1 limits1 and definitions of the species into which he groups the very same things4 ,in these islands1 we are in the habit of regarding mankind as of one species1 but a fortnight's steam will land us in a country where divines and savants1 for once in agreement1 vie with one another in loudness of assertion1 if not in cogency of proof1 that men are of different species2 and1 more particularly1 that the species negro is so distinct from our own that the ,ten ,commandments have actually no reference to him4 ,even in the calm region of entomology1 where1 if anywhere in this sinful world1 passion and prejudice should fail to stir the mind1 one learned coleopterist will fill ten attractive volumes with descriptions of species of beetles1 nine-tenths of which are immediately declared by his brother beetle-mongers to be no species at all4 ,the truth is that the number of distinguishable living creatures almost surpasses imagination4 ,at least a hundred thousand such kinds of insects alone have been described and may be identified in collections1 and the number of separable kinds of living things is underestimated at half a million4 ,seeing that most of these obvious kinds have their accidental varieties1 and that they often shade into others by imperceptible degrees1 it may well be imagined that the task of distinguishing between what is permanent and what fleeting1 what is a species and what a mere variety1 is sufficiently formidable4 ,but is it not possible to apply a test whereby a true species may be known from a mere variety8 ,is there no criterion of species8 ,great authorities affirm that there is--that the unions of members of the same species are always fertile1 while those of distinct species are either sterile1 or their offspring1 called hybrids1 are so4 ,it is affirmed not only that this is an experimental fact1 but that it is a provision for the preservation of the purity of species4 ,such a criterion as this would be invaluable2 but1 unfortunately1 not only is it not obvious how to apply it in the great majority of cases in which its aid is needed1 but its general validity is stoutly denied4 ,the ,hon4 and ,rev4 ,mr4 ,herbert1 a most trustworthy authority1 not only asserts as the result of his own observations and experiments that many hybrids are quite as fertile as the parent species1 but he goes so far as to assert that the particular plant ,crinum capense is much more fertile when crossed by a distinct species than when fertilised by its proper pollen6 ,on the other hand1 the famous ,gaertner1 though he took the greatest pains to cross the primrose and the cowslip1 succeeded only once or twice in several years2 and yet it is a well-established fact that the primrose and the cowslip are only varieties of the same kind of plant4 ,again1 such cases as the following are well established4 ,the female of species ,a1 if crossed with the male of species ,b1 is fertile2 but1 if the female of ,b is crossed with the male of ,a1 she remains barren4 ,facts of this kind destroy the value of the supposed criterion4 ,if1 weary of the endless difficulties involved in the determination of species1 the investigator1 contenting himself with the rough practical distinction of separable kinds1 endeavours to study them as they occur in nature--to ascertain their relations to the conditions which surround them1 their mutual harmonies and discordances of structure1 the bond of union of their parts and their past history1 he finds himself1 according to the received notions1 in a mighty maze1 and with1 at most1 the dimmest adumbration of a plan4 ,if he starts with any one clear conviction1 it is that every part of a living creature is cunningly adapted to some special use in its life4 ,has not his ,paley told him that that seemingly useless organ1 the spleen1 is beautifully adjusted as so much packing between the other organs8 ,and yet1 at the outset of his studies1 he finds that no adaptive reason whatsoever can be given for one-half of the peculiarities of vegetable structure2 he also discovers rudimentary teeth1 which are never used1 in the gums of the young calf and in those of the foetal whale2 insects which never bite have rudimental jaws1 and others which never fly have rudimental wings2 naturally blind creatures have rudimental eyes2 and the halt have rudimentary limbs4 ,so1 again1 no animal or plant puts on its perfect form at once1 but all have to start from the same point1 however various the course which each has to pursue4 ,not only men and horses1 and cats and dogs1 lobsters and beetles1 periwinkles and mussels1 but even the very sponges and animalcules commence their existence under forms which are essentially undistinguishable2 and this is true of all the infinite variety of plants4 ,nay1 more1 all living beings march side by side along the high road of development1 and separate the later the more like they are2 like people leaving church1 who all go down the aisle1 but having reached the door some turn into the parsonage1 others go down the village1 and others part only in the next parish4 ,a man in his development runs for a little while parallel with1 though never passing through1 the form of the meanest worm1 then travels for a space beside the fish1 then journeys along with the bird and the reptile for his fellow travellers2 and only at last1 after a brief companionship with the highest of the four-footed and four-handed world1 rises into the dignity of pure manhood4 ,no competent thinker of the present day dreams of explaining these indubitable facts by the notion of the existence of unknown and undiscoverable adaptations to purpose4 ,and we would remind those who1 ignorant of the facts1 must be moved by authority1 that no one has asserted the incompetence of the doctrine of final causes1 in its application to physiology and anatomy1 more strongly than our own eminent anatomist1 ,professor ,owen1 who1 speaking of such cases1 says 7,on the ,nature of ,limbs1 pp4 #ci1 #dj73 8,i think it will be obvious that the principle of final adaptations fails to satisfy all the conditions of the problem40 ,but1 if the doctrine of final causes will not help us to comprehend the anomalies of living structure1 the principle of adaptation must surely lead us to understand why certain living beings are found in certain regions of the world and not in others4 ,the palm1 as we know1 will not grow in our climate1 nor the oak in ,greenland4 ,the white bear cannot live where the tiger thrives1 nor vice versa1 and the more the natural habits of animal and vegetable species are examined1 the more do they seem1 on the whole1 limited to particular provinces4 ,but when we look into the facts established by the study of the geographical distribution of animals and plants it seems utterly hopeless to attempt to understand the strange and apparently capricious relations which they exhibit4 ,one would be inclined to suppose a priori that every country must be naturally peopled by those animals that are fittest to live and thrive in it4 ,and yet how1 on this hypothesis1 are we to account for the absence of cattle in the ,pampas of ,south ,america1 when those parts of the ,new ,world were discovered8 ,it is not that they were unfit for cattle1 for millions of cattle now run wild there2 and the like holds good of ,australia and ,new ,zealand4 ,it is a curious circumstance1 in fact1 that the animals and plants of the ,northern ,hemisphere are not only as well adapted to live in the ,southern ,hemisphere as its own autochthones1 but are in many cases absolutely better adapted1 and so overrun and extirpate the aborigines4 ,clearly1 therefore1 the species which naturally inhabit a country are not necessarily the best adapted to its climate and other conditions4 ,the inhabitants of islands are often distinct from any other known species of animal or plants 7witness our recent examples from the work of ,sir ,emerson ,tennent1 on ,ceylon71 and yet they have almost always a sort of general family resemblance to the animals and plants of the nearest mainland4 ,on the other hand1 there is hardly a species of fish1 shell1 or crab common to the opposite sides of the narrow isthmus of ,panama4 ,wherever we look1 then1 living nature offers us riddles of difficult solution1 if we suppose that what we see is all that can be known of it4 ,but our knowledge of life is not confined to the existing world4 ,whatever their minor differences1 geologists are agreed as to the vast thickness of the accumulated strata which compose the visible part of our earth1 and the inconceivable immensity of the time of whose lapse they are the imperfect1 but the only accessible witnesses4 ,now1 throughout the greater part of this long series of stratified rocks are scattered1 sometimes very abundantly1 multitudes of organic remains1 the fossilized exuviae of animals and plants which lived and died while the mud of which the rocks are formed was yet soft ooze1 and could receive and bury them4 ,it would be a great error to suppose that these organic remains were fragmentary relics4 ,our museums exhibit fossil shells of immeasurable antiquity1 as perfect as the day they were formed1 whole skeletons without a limb disturbed--nay1 the changed flesh1 the developing embryos1 and even the very footsteps of primeval organisms4 ,thus the naturalist finds in the bowels of the earth species as well defined as1 and in some groups of animals more numerous than1 those that breathe the upper air4 ,but1 singularly enough1 the majority of these entombed species are wholly distinct from those that now live4 ,nor is this unlikeness without its rule and order4 ,as a broad fact1 the further we go back in time the less the buried species are like existing forms2 and the further apart the sets of extinct creatures are the less they are like one another4 ,in other words1 there has been a regular succession of living beings1 each younger set being in a very broad and general sense somewhat more like those which now live4 ,it was once supposed that this succession had been the result of vast successive catastrophes1 destructions1 and re-creations en masse2 but catastrophes are now almost eliminated from geological1 or at least palaeontological speculation2 and it is admitted on all hands that the seeming breaks in the chain of being are not absolute1 but only relative to our imperfect knowledge2 that species have replaced species1 not in assemblages1 but one by one2 and that1 if it were possible to have all the phenomena of the past presented to us1 the convenient epochs and formations of the geologist1 though having a certain distinctness1 would fade into one another with limits as undefinable as those of the distinct and yet separable colours of the solar spectrum4 ,such is a brief summary of the main truths which have been established concerning species4 ,are these truths ultimate and irresolvable facts1 or are their complexities and perplexities the mere expressions of a higher law8 ,a large number of persons practically assume the former position to be correct4 ,they believe that the writer of the ,pentateuch was empowered and commissioned to teach us scientific as well as other truth1 that the account we find there of the creation of living things is simply and literally correct1 and that anything which seems to contradict it is1 by the nature of the case1 false4 ,all the phenomena which have been detailed are1 on this view1 the immediate product of a creative fiat and consequently are out of the domain of science altogether4 ,whether this view prove ultimately to be true or false1 it is1 at any rate1 not at present supported by what is commonly regarded as logical proof1 even if it be capable of discussion by reason2 and hence we consider ourselves at liberty to pass it by1 and to turn to those views which profess to rest on a scientific basis only1 and therefore admit of being argued to their consequences4 ,and we do this with the less hesitation as it so happens that those persons who are practically conversant with the facts of the case 7plainly a considerable advantage7 have always thought fit to range themselves under the latter category4 ,the majority of these competent persons have up to the present time maintained two positions1--the first1 that every species is1 within certain defined or definable limits1 fixed and incapable of modification2 the second1 that every species was originally produced by a distinct creative act4 ,the second position is obviously incapable of proof or disproof1 the direct operations of the ,creator not being subjects of science2 and it must therefore be regarded as a corollary from the first1 the truth or falsehood of which is a matter of evidence4 ,most persons imagine that the arguments in favour of it are overwhelming2 but to some few minds1 and these1 it must be confessed1 intellects of no small power and grasp of knowledge1 they have not brought conviction4 ,among these minds1 that of the famous naturalist ,lamarck1 who possessed a greater acquaintance with the lower forms of life than any man of his day1 ,cuvier not excepted1 and was a good botanist to boot1 occupies a prominent place4 ,two facts appear to have strongly affected the course of thought of this remarkable man--the one1 that finer or stronger links of affinity connect all living beings with one another1 and that thus the highest creature grades by multitudinous steps into the lowest2 the other1 that an organ may be developed in particular directions by exerting itself in particular ways1 and that modifications once induced may be transmitted and become hereditary4 ,putting these facts together1 ,lamarck endeavoured to account for the first by the operation of the second4 ,place an animal in new circumstances1 says he1 and its needs will be altered2 the new needs will create new desires1 and the attempt to gratify such desires will result in an appropriate modification of the organs exerted4 ,make a man a blacksmith1 and his brachial muscles will develop in accordance with the demands made upon them1 and in like manner1 says ,lamarck1 8the efforts of some short-necked bird to catch fish without wetting himself have1 with time and perseverance1 given rise to all our herons and long-necked waders40 ,the ,lamarckian hypothesis has long since been justly condemned1 and it is the established practice for every tyro to raise his heel against the carcass of the dead lion4 ,but it is rarely either wise or instructive to treat even the errors of a really great man with mere ridicule1 and in the present case the logical form of the doctrine stands on a very different footing from its substance4 ,if species have really arisen by the operation of natural conditions1 we ought to be able to find those conditions now at work2 we ought to be able to discover in nature some power adequate to modify any given kind of animal or plant in such a manner as to give rise to another kind1 which would be admitted by naturalists as a distinct species4 ,lamarck imagined that he had discovered this vera causa in the admitted facts that some organs may be modified by exercise2 and that modifications1 once produced1 are capable of hereditary transmission4 ,it does not seem to have occurred to him to inquire whether there is any reason to believe that there are any limits to the amount of modification producible1 or to ask how long an animal is likely to endeavour to gratify an impossible desire4 ,the bird1 in our example1 would surely have renounced fish dinners long before it had produced the least effect on leg or neck4 ,since ,lamarck's time1 almost all competent naturalists have left speculations on the origin of species to such dreamers as the author of the 8,vestiges10 by whose well-intentioned efforts the ,lamarckian theory received its final condemnation in the minds of all sound thinkers4 ,notwithstanding this silence1 however1 the transmutation theory1 as it has been called1 has been a 8skeleton in the closet0 to many an honest zoologist and botanist who had a soul above the mere naming of dried plants and skins4 ,surely1 has such an one thought1 nature is a mighty and consistent whole1 and the providential order established in the world of life must1 if we could only see it rightly1 be consistent with that dominant over the multiform shapes of brute matter4 ,but what is the history of astronomy1 of all the branches of physics1 of chemistry1 of medicine1 but a narration of the steps by which the human mind has been compelled1 often sorely against its will1 to recognize the operation of secondary causes in events where ignorance beheld an immediate intervention of a higher power8 ,and when we know that living things are formed of the same elements as the inorganic world1 that they act and react upon it1 bound by a thousand ties of natural piety1 is it probable1 nay is it possible1 that they1 and they alone1 should have no order in their seeming disorder1 no unity in their seeming multiplicity1 should suffer no explanation by the discovery of some central and sublime law of mutual connexion8 ,questions of this kind have assuredly often arisen1 but it might have been long before they received such expression as would have commanded the respect and attention of the scientific world1 had it not been for the publication of the work which prompted this article4 ,its author1 ,mr4 ,darwin1 inheritor of a once celebrated name1 won his spurs in science when most of those now distinguished were young men1 and has for the last #bj years held a place in the front ranks of ,british philosophers4 ,after a circumnavigatory voyage1 undertaken solely for the love of his science1 ,mr4 ,darwin published a series of researches which at once arrested the attention of naturalists and geologists2 his generalizations have since received ample confirmation1 and now command universal assent1 nor is it questionable that they have had the most important influence on the progress of science4 ,more recently ,mr4 ,darwin1 with a versatility which is among the rarest of gifts1 turned his attention to a most difficult question of zoology and minute anatomy2 and no living naturalist and anatomist has published a better monograph than that which resulted from his labours4 ,such a man1 at all events1 has not entered the sanctuary with unwashed hands1 and when he lays before us the results of #bj years' investigation and reflection we must listen even though we be disposed to strike4 ,but1 in reading his work it must be confessed that the attention which might at first be dutifully1 soon becomes willingly1 given1 so clear is the author's thought1 so outspoken his conviction1 so honest and fair the candid expression of his doubts4 ,those who would judge the book must read it2 we shall endeavour only to make its line of argument and its philosophical position intelligible to the general reader in our own way4 ,the ,baker-street ,bazaar has just been exhibiting its familiar annual spectacle4 ,straight-backed1 small-headed1 big-barrelled oxen1 as dissimilar from any wild species as can well be imagined1 contended for attention and praise with sheep of half-a-dozen different breeds and styes of bloated preposterous pigs1 no more like a wild boar or sow than a city alderman is like an ourang-outang4 ,the cattle show has been1 and perhaps may again be1 succeeded by a poultry show1 of whose crowing and clucking prodigies it can only be certainly predicated that they will be very unlike the aboriginal ,phasianus gallus4 ,if the seeker after animal anomalies is not satisfied1 a turn or two in ,seven ,dials will convince him that the breeds of pigeons are quite as extraordinary and unlike one another and their parent stock1 while the ,horticultural ,society will provide him with any number of corresponding vegetable aberrations from nature's types4 ,he will learn with no little surprise1 too1 in the course of his travels1 that the proprietors and producers of these animal and vegetable anomalies regard them as distinct species1 with a firm belief1 the strength of which is exactly proportioned to their ignorance of scientific biology1 and which is the more remarkable as they are all proud of their skill in originating such 8species40 ,on careful inquiry it is found that all these1 and the many other artificial breeds or races of animals and plants1 have been produced by one method4 ,the breeder--and a skilful one must be a person of much sagacity and natural or acquired perceptive faculty--notes some slight difference1 arising he knows not how1 in some individuals of his stock4 ,if he wish to perpetuate the difference1 to form a breed with the peculiarity in question strongly marked1 he selects such male and female individuals as exhibit the desired character1 and breeds from them4 ,their offspring are then carefully examined1 and those which exhibit the peculiarity the most distinctly are selected for breeding1 and this operation is repeated until the desired amount of divergence from the primitive stock is reached4 ,it is then found that by continuing the process of selection--always breeding1 that is1 from well-marked forms1 and allowing no impure crosses to interfere1--a race may be formed1 the tendency of which to reproduce itself is exceedingly strong2 nor is the limit to the amount of divergence which may be thus produced known1 but one thing is certain1 that1 if certain breeds of dogs1 or of pigeons1 or of horses1 were known only in a fossil state1 no naturalist would hesitate in regarding them as distinct species4 ,but1 in all these cases we have human interference4 ,without the breeder there would be no selection1 and without the selection no race4 ,before admitting the possibility of natural species having originated in any similar way1 it must be proved that there is in nature some power which takes the place of man1 and performs a selection sua sponte4 ,it is the claim of ,mr4 ,darwin that he professes to have discovered the existence and the modus operandi of this natural selection1 as he terms it2 and1 if he be right1 the process is perfectly simple and comprehensible1 and irresistibly deducible from very familiar but well nigh forgotten facts4 ,who1 for instance1 has duly reflected upon all the consequences of the marvellous struggle for existence which is daily and hourly going on among living beings8 ,not only does every animal live at the expense of some other animal or plant1 but the very plants are at war4 ,the ground is full of seeds that cannot rise into seedlings2 the seedlings rob one another of air1 light and water1 the strongest robber winning the day1 and extinguishing his competitors4 ,year after year1 the wild animals with which man never interferes are1 on the average1 neither more nor less numerous than they were2 and yet we know that the annual produce of every pair is from one to perhaps a million young1--so that it is mathematically certain that1 on the average1 as many are killed by natural causes as are born every year1 and those only escape which happen to be a little better fitted to resist destruction than those which die4 ,the individuals of a species are like the crew of a foundered ship1 and none but good swimmers have a chance of reaching the land4 ,such being unquestionably the necessary conditions under which living creatures exist1 ,mr4 ,darwin discovers in them the instrument of natural selection4 ,suppose that in the midst of this incessant competition some individuals of a species 7,a7 present accidental variations which happen to fit them a little better than their fellows for the struggle in which they are engaged1 then the chances are in favour1 not only of these individuals being better nourished than the others1 but of their predominating over their fellows in other ways1 and of having a better chance of leaving offspring1 which will of course tend to reproduce the peculiarities of their parents4 ,their offspring will1 by a parity of reasoning1 tend to predominate over their contemporaries1 and there being 7suppose7 no room for more than one species such as ,a1 the weaker variety will eventually be destroyed by the new destructive influence which is thrown into the scale1 and the stronger will take its place4 ,surrounding conditions remaining unchanged1 the new variety 7which we may call ,b7--supposed1 for argument's sake1 to be the best adapted for these conditions which can be got out of the original stock--will remain unchanged1 all accidental deviations from the type becoming at once extinguished1 as less fit for their post than ,b itself4 ,the tendency of ,b to persist will grow with its persistence through successive generations1 and it will acquire all the characters of a new species4 ,but1 on the other hand1 if the conditions of life change in any degree1 however slight1 ,b may no longer be that form which is best adapted to withstand their destructive1 and profit by their sustaining1 influence2 in which case if it should give rise to a more competent variety 7,c71 this will take its place and become a new species2 and thus1 by natural selection1 the species ,b and ,c will be successively derived from ,a4 ,that this most ingenious hypothesis enables us to give a reason for many apparent anomalies in the distribution of living beings in time and space1 and that it is not contradicted by the main phenomena of life and organization appear to us to be unquestionable2 and so far it must be admitted to have an immense advantage over any of its predecessors4 ,but it is quite another matter to affirm absolutely either the truth or falsehood of ,mr4 ,darwin's views at the present stage of the inquiry4 ,goethe has an excellent aphorism defining that state of mind which he calls ,thtige ,skepsis--active doubt4 ,it is doubt which so loves truth that it neither dares rest in doubting1 nor extinguish itself by unjustified belief2 and we commend this state of mind to students of species1 with respect to ,mr4 ,darwin's or any other hypothesis1 as to their origin4 ,the combined investigations of another #bj years may1 perhaps1 enable naturalists to say whether the modifying causes and the selective power1 which ,mr4 ,darwin has satisfactorily shown to exist in nature1 are competent to produce all the effects he ascribes to them1 or whether1 on the other hand1 he has been led to over-estimate the value of his principle of natural selection1 as greatly as ,lamarck overestimated his vera causa of modification by exercise4 ,but there is1 at all events1 one advantage possessed by the more recent writer over his predecessor4 ,mr4 ,darwin abhors mere speculation as nature abhors a vacuum4 ,he is as greedy of cases and precedents as any constitutional lawyer1 and all the principles he lays down are capable of being brought to the test of observation and experiment4 ,the path he bids us follow professes to be1 not a mere airy track1 fabricated of ideal cobwebs1 but a solid and broad bridge of facts4 ,if it be so1 it will carry us safely over many a chasm in our knowledge1 and lead us to a region free from the snares of those fascinating but barren ,virgins1 the ,final ,causes1 against whom a high authority has so justly warned us4 8,my sons1 dig in the vineyard10 were the last words of the old man in the fable2 and1 though the sons found no treasure1 they made their fortunes by the grapes4 99 ,times1 ,december #bfth1 #ahej4